Collodictyonidae

Collodictyonidae
	Collodictyon sp.
Scientific classification
Domain:	Eukaryota
Clade:	Opimoda
Clade:	Podiata
Clade:	CRuMs
Family:	Collodictyonidae; Brugerolle et al. 2002
Genera
	Collodictyon Carter 1865; Diphylleia Massart 1920 non Michaux 1803;
Synonyms
	Diphylleidae Cavalier-Smith 1993;

Collodictyonidae (also Diphylleidae) is a group of aquatic, unicellular eukaryotic organisms with two to four terminal flagella. They feed by phagocytosis, ingesting other unicellular organisms like algae and bacteria. The most remarkable fact of this clade is its uncertain position in the tree of life. A recent phylogenomic analysis places it either as sister group of the excavate protist Malawimonas or as sister group of the Bikonta clade, although both positions received considerably low branch support values.[1] This means that Collodictyonidae is somewhere near the root of the Eukaryote tree, probably close to the unikont–bikont bifurcation.

Recent molecular analyses place Collodictyonids (e.g. Collodictyon) ins a clade also containing Rigifilida and Mantamonadidae. This clade has been name CRuMs and is sister to Amorphea.[2][3]

References

Zhao, Sen; Fabien Burki; Jon Bråte; Patrick Keeling; Dag Klaveness; Kamran Shalchian-Tabrizi (2012). "Collodictyon – An Ancient Lineage in the Tree of Eukaryotes". Molecular Biology and Evolution. 29 (6): 1557–68. doi:10.1093/molbev/mss001. PMC 3351787. PMID 22319147.
Lax, Gordon; Eglit, Yana; Eme, Laura; Bertrand, Erin M.; Roger, Andrew J.; Simpson, Alastair G. B. (2018-11-14). "Hemimastigophora is a novel supra-kingdom-level lineage of eukaryotes". Nature. 564 (7736): 410–414. doi:10.1038/s41586-018-0708-8. ISSN 0028-0836. PMID 30429611.
Brown, Matthew W; Heiss, Aaron A; Kamikawa, Ryoma; Inagaki, Yuji; Yabuki, Akinori; Tice, Alexander K; Shiratori, Takashi; Ishida, Ken-Ichiro; Hashimoto, Tetsuo (2018-01-19). "Phylogenomics Places Orphan Protistan Lineages in a Novel Eukaryotic Super-Group". Genome Biology and Evolution. 10 (2): 427–433. doi:10.1093/gbe/evy014. ISSN 1759-6653. PMC 5793813. PMID 29360967.

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[1] Zhao, Sen; Fabien Burki; Jon Bråte; Patrick Keeling; Dag Klaveness; Kamran Shalchian-Tabrizi (2012). "Collodictyon – An Ancient Lineage in the Tree of Eukaryotes". Molecular Biology and Evolution. 29 (6): 1557–68. doi:10.1093/molbev/mss001. PMC 3351787. PMID 22319147.

[2] Lax, Gordon; Eglit, Yana; Eme, Laura; Bertrand, Erin M.; Roger, Andrew J.; Simpson, Alastair G. B. (2018-11-14). "Hemimastigophora is a novel supra-kingdom-level lineage of eukaryotes". Nature. 564 (7736): 410–414. doi:10.1038/s41586-018-0708-8. ISSN 0028-0836. PMID 30429611.

[3] Brown, Matthew W; Heiss, Aaron A; Kamikawa, Ryoma; Inagaki, Yuji; Yabuki, Akinori; Tice, Alexander K; Shiratori, Takashi; Ishida, Ken-Ichiro; Hashimoto, Tetsuo (2018-01-19). "Phylogenomics Places Orphan Protistan Lineages in a Novel Eukaryotic Super-Group". Genome Biology and Evolution. 10 (2): 427–433. doi:10.1093/gbe/evy014. ISSN 1759-6653. PMC 5793813. PMID 29360967.